The Nittany Lions were busy following the Orange Bowl. Miss any roster changes? Catch up here.

A flurry of roster moves last week has provided a much clearer picture of what Penn State football projects to look like in 2025. With many upperclassmen returning, the Nittany Lions’ “job’s not finished” mentality could propel them further than the semifinals. However, question marks at key positions also could hold them back from reaching the championship.

The news broke constantly last week, so here’s what Penn State looks like after a frantic pace of post-Orange Bowl change.

Penn State’s run game, offensive line look monstrous

With Nick Singleton and Kaytron Allen returning for their fourth years, Penn State could have the best backfield in the nation. Singleton and Allen both had career highs in rushing yards last season as the program’s first 1,000-yard rushing duo. With another year in offensive coordinator Andy Kotelnicki’s system, these two could be even more special. It’s also worth noting that both made strides as receiving backs last year, which could be very relevant given Penn State’s lack of proven receiving targets on the roster.

The offensive line was a big part of the team’s success last year and has lost only one starter to the NFL in guard Sal Wormley. Cooper Cousins, who burned his redshirt last season, is the favorite to start at guard after earning rave reviews from coaches and players in his first year with the team. Veteran JB Nelson could also start. Both players are great utility guys on the line and should see playing time somewhere on the line next year.

Tackle Drew Shelton, guard Vega Ioane and center Nick Dawkins are expected to remain in their starting roles next year, leaving the only other question mark at right tackle. Anthony Donkoh was excellent as the starter last season, but after he went down, Nolan Rucci did a great job in his place. Depending on Donkoh’s recovery timeline, it could be Rucci getting the nod at right tackle to start the season.